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Disclosure - Research Collaboration and License Agreements link:presentationLink link:calculationLink link:definitionLink 2322304 - Disclosure - Research Collaboration and License Agreements (Tables) link:presentationLink link:calculationLink link:definitionLink 2423413 - Disclosure - Research Collaboration and License Agreements - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2424414 - Disclosure - Research Collaboration and License Agreements - Schedule of Milestone Payments by Indications (Details) link:presentationLink link:calculationLink link:definitionLink 2425415 - Disclosure - Research Collaboration and License Agreements - Schedule of Recognized Revenue from Collaboration and License Agreements (Details) link:presentationLink link:calculationLink link:definitionLink 2126106 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 2327305 - Disclosure - Commitments and Contingencies (Tables) link:presentationLink link:calculationLink link:definitionLink 2428416 - Disclosure - Commitments and Contingencies - Additional Information (Details) link:presentationLink link:calculationLink link:definitionLink 2429417 - Disclosure - Commitments and Contingencies - Components of Lease Costs (Details) link:presentationLink link:calculationLink link:definitionLink 2430418 - Disclosure - Commitments and Contingencies - Operating Lease Liability Maturity (Details) link:presentationLink link:calculationLink link:definitionLink 2430418 - Disclosure - Commitments and Contingencies - Operating Lease Liability Maturity (Details) link:presentationLink link:calculationLink link:definitionLink 2431419 - Disclosure - Commitments and Contingencies - Future Minimum Commitments under Unassigned Lease Obligations (Details) link:presentationLink link:calculationLink link:definitionLink 2132107 - Disclosure - Stockholders' Equity link:presentationLink link:calculationLink link:definitionLink 2333306 - Disclosure - Stockholders' Equity (Tables) link:presentationLink link:calculationLink link:definitionLink 2434420 - 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